Sr2 is a durable stem rust resistance gene used in rust-prone areas and has been a significant source in the global control of this pathogen. Sr2 is closely associated with Lr27, Yr30 and with pseudo-black chaff which involves melanin pigmentation of the glumes and stem, particularly below the uppermost node. It was introgressed from T. turgidum var. dicoccum cv. Yaroslav into bread wheat by McFadden in 1930.